Wednesday, December 30, 2009

The Maltese letter 'għ' in taxonomy

The 9th letter of the Maltese alphabet is a digraph written as and pronounced 'għajn' (aighn in English). In speech, it is a silent consonant which has the function of lengthening the succeeding vowel, therefore a word written as 'għar' (meaning 'cave') would be pronounced something like 'aar'.

Several placenames in the Maltese Islands bear the letter għ - Għarb, Bengħisa and Għargħur being a few examples. Taxonomists naming animals from such places for their type locality have to use gh instead of in the specific epithet, simply because the ICZN does not allow for any characters other than the 26 of the Latin alphabet (Article 11.2).

Through the literature cited, I was able to come across five such animals described from the Maltese Islands. Clio ghawdexensis Janssen, 2003 is a fossil pteropod named after the island of Gozo, known in Maltese as Għawdex. Trochoidea gharlapsi Beckmann, 1987 is another snail, this time a recent terrestrial species localized mainly around the area of Għar Lapsi ('Lapsi Cave') in southern Malta (juvenile specimen illustrated below, photographed in December 2009).

However, both the above-mentioned type localities are (nomenclaturally) one-offs. The place to boast most species named for it is Għar Dalam (Dalam Cave), which is an extremely important location for fossils of the dwarf elephants and hippopotami that roamed Malta during the Pleistocene. Armadillidium ghardalamensis Caruso & Hili, 1991 is a woodlouse that is endemic to this cave and another one close by (Għar Ħassan). Scolopax ghardalamensis Fischer & Stephan, 1974 is a fossil bird described from the place, though this is now known to be a junior synonym of Coturnix coturnix (Linn., 1758). Finally, Myotis ghardalamensis Storch, 1974, an early Pleistocene bat, probably ancestral to the more modern species.

The million-dollar question is, how would these names, purely hypothetically or actually, be pronounced? Would the Maltese letter retain or lose its linguistic characteristics when included in a species' Latinized name? If binomial names were commonly used, would one speak of Trochoidea 'garlapsi' [sic], or Trochoidea 'aarlapsi' [sic]?

References:


Beckmann, K. -H. (1987). Land und Süßwassermollusken der Maltesischen Inseln. Heldia, 1 (Sonderheft): 1-38.
Caruso, D. & Hili, C. (1991). Nuovi dati sugli isopodi terrestri delle isole dell’arcipelago maltese. Animalia, 18: 115-124 .
Fischer, K. von & Stephan, B. (1974). Eine pleistozäne Avifauna aus der Ghar Dalam-Höhle, Malta. Zeitschrift für Geologische Wissenschaften, 2 (4): 515-523.
Janssen, A. W. (2004), Fossils from the Lower Globigerina Limestone Formation at Wardija, Gozo (Miocene, Aquitanian), with a description of some new pteropod species (Mollusca, Gastropoda). The Central Mediterranean Naturalist, 4 (1): 1-33.
Olsson, S. L. (1976). Fossil woodcocks: an extinct species from Puerto Rico and an invalid species from Malta (Aves: Scolopacidae: Scolopax). Proceedings of the Biological Society of Washington, 89 (20): 265-274.
Storch, G. (1974): Quartäre Fledermaus-Faunen von der Insel Malta. Senckenbergiana Lethaea, 55: 407-434.

Note: I edited my own comment below to articulate a sentence better, the original comment has been deleted and replaced with the slightly amended one.

Friday, December 25, 2009

Melarhaphe neritoides - before and after

Melarhaphe neritoides (Linn., 1758) is a seasnail which spends the summer months aestivating in cracks on bare rock, in a part of the shore where the sea does not reach, sealing up its aperture with a solid horny operculum to prevent losing water in the extreme heat accumulating during the day.

As soon as winter comes, more powerful waves and heavier mist start wetting the area where the snail lives, eventually filling pools in the rock and allowing the snail to resume its 'normal' lifestyle - moving about, grazing on algae, mating and laying eggs, in time before the dryness returns.

The following photographs were taken in the same rock hollow in August (photo 1) and December (photo 2) on the limestone shore of Buġibba, Malta, showing the snail during its period of inactivity and its subsequent 'resurrection'.




Tuesday, December 15, 2009

Research Paper 2

Cilia, D. P. (2009). On the presence of the alien freshwater gastropod Ferrissia fragilis (Tryon, 1863) (Gastropoda: Planorbidae) in the Maltese Islands (Central Mediterranean). Bollettino Malacologico, 45: 123-127.

Downloadable here: http://www.box.net/shared/qpfhcslgpf